Why does keyboard shortcut using alt-xx beep?

Whenever I use the keyboard shortcuts with an alt prefix (e.g. alt-F, alt-C, alt-B, alt-S, alt-G) in the offline editor, the system beeps as though it were an error.  Everything seems to work correctly otherwise.  Why is it beeping?  It doesn't do it if I use the virtual keyboard, only if I use the keyboard shortcuts. Is there a way to stop this?  It seems to happen with every shortcut the uses alt-xx except for alt-. (Fan) that was recently added. 

Related